It is the application of the colors, sometimes a bit sloppy and sometimes quite precise, together with the state of preservation of the miniatures, that creates difficulties for modern viewers.
It is helpful to keep in mind some of the possible causes, whether technical or due to the differences of more than one engaged colorist, as to why the painting process is so uneven.
From: “Reconceptualizing the Poems of the Pearl-Gawain Manuscript in Line and Color,” Manuscript Studies 3.2 (2018), 383-420.
The present study will both explore further the reconceptualization of the poems (in MS Cotton Nero A.x) by...
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The implications of the new scientific data, which Paul Garside, the Conservation Scientist at the British Library, provided in answer to my queries regarding the pigments, will be examined here.
His "Analysis of Pigments found in Cotton Nero A.x" is available in my Appendix (pages 127-31).
One of the major discoveries is that the ink of the underdrawings and the text, as I had anticipated, is the same.
Further, in the scientifically enhanced images, it is now possible to see how the added paint of the miniatures obscured visual information not previously apprehended.